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首页> 外文期刊>IAWA Journal >Structurally complex, yet anatomically plesiomorphic: permineralized plants from the Emsian of Gaspe (Quebec, Canada) expand the diversity of Early Devonian euphyllophytes
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Structurally complex, yet anatomically plesiomorphic: permineralized plants from the Emsian of Gaspe (Quebec, Canada) expand the diversity of Early Devonian euphyllophytes

机译：结构复杂，但解剖学性质：来自汽油Emsian（加拿大魁北克，加拿大）的渗透性植物扩大了早期牧民野外卵骨菌素的多样性

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The Battery Point Formation of eastern Canada hosts an Emsian (c. 400-395 Ma) flora that marks one of the rare occurrences of anatomically-preserved Early Devonian plants. We describe four new euphyllophytes from small permineralized axes in this unit. Leptocentroxyla tetrarcha gen. et sp. nov. has a four-ribbed mesarch actinostele with Psilophyton-type (P-type) tracheids and a central area of scalariform tracheids. Stenoloboxyla ambigua gen. et sp. nov. has a bar-shaped to three-ribbed mesarch stele lacking central protoxylem, with one of the ribs less pronounced, P-type tracheids, and sclerenchyma forming a discontinuous layer in the cortex. Jowingera triloba gen. et sp. nov. has a three-ribbed mesarch actinostele with central protoxylem and P-type tracheids. Tainioxyla quebecana gen. et sp. nov. has bar-shaped xylem with mesarch protoxylem strands, P-type tracheids, and anatomy typical of cambial growth initiation. These new species raise the diversity of Battery Point Formation permineralized plants to nine genera, adding significantly to the diversity of Early Devonian plants characterized anatomically. The four species encompass structural diversity of unexpected breadth and novelty for their age. They are different from both older and coeval euphyllophytes and from younger euphyllophytes, exhibiting combinations of derived and plesiomorphic characters. Their mesarch actinosteles and bar-shaped protosteles, histological differentiation within metaxylem and cortex, and secondary growth, represent aspects of structural complexity common in more derived Middle-Late Devonian euphyllophytes. Concurrently, the four species share P-type tracheids typical of Early Devonian basal euphyllophytes with simpler anatomies. These new fossils offer a first glimpse of a plexus of plants representing a previously unsuspected stage of euphyllophyte morphoanatomical evolution. They demonstrate significant euphyllophyte diversification and exploration of structural complexity under way during the Early Devonian, against a background of plesiomorphic-type tracheids. When more completely characterized, these Emsian plants will provide links for resolving phylogenetic relationships at the base of the euphyllophyte clade.
机译：加拿大东部的电池点形成托管EMSIAN（C.400-395 MA）植物群，标志着解剖学保存的早期德文植物的罕见发生之一。我们在本机中描述了来自小型渗透轴的四种新的野外野外胰岛素。 Leptocentroxyla Tetrarcha Gen。等等。 11月。有一个四罗纹的Mesarch actinostele，具有psilophyton型（p型）躯干和脉管中的中心区域。 Stenoloboxyla Ambigua Gen。等等。 11月。有一个棒状到三罗纹椎管架缺乏中央protoxylem，其中一个肋骨较小，p型躯干，并且硬化在皮质中形成不连续层。 Jowingera Triloba Gen。等等。 11月。有一个三罗纹的mesarch actinostele，中央protoxylem和p型肱骨。 tainioxyla quebecana gen。等等。 11月。具有杆状木质丝绒，具有胶质淀粉鳞片，p型状鳞片，以及冠状生长引发的解剖学。这些新物种提高了电池点形成型渗透性植物的多样性，达到九个世纪，对解剖学表征的早期牧民植物的多样性显着增加。这四种物种包括出乎意料的广度和新颖性的结构多样性。它们与较旧的卵骨细胞和来自较年轻的野外野生植物不同，表现出衍生和衍生和渗透性的组合。它们的Mesarch Actinosteles和棒状致辐射，组织学分化，组织学分化和皮质和皮质和二次生长，代表了在更衍生的中期德文郡嗜酸盐卵体中常见的结构复杂性的方面。同时，四种物种共享P型Trachids典型的早期牧民基础野外野外卵囊细胞，具有更简单的解剖。这些新化石提供了代表euphylophyte Morphoanatomical演进的先前未缺乏阶段的植物丛丛中的第一眼。他们在早期德文郡在普通晶型治疗后的背景下表明了在早期德文郡的术后结构复杂性的显着野外嗜植物多样化和探索。当更完全表征时，这些EMSian植物将提供用于解决euphyllophyte植物碱在野外发育关系的链接。

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